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Two crystal structures of a plant endo-1,3-β-glucanase representing glycoside hydrolase family GH17 in complexes with products of laminarahexose hydrolysis reveal the protein–saccharide interactions at five key binding subsites for the first time. A trisaccharide is bound at subsites −3, −2 and −1 of the nonreducing end (glycon portion) in both structures, and in one structure an extra tetrasaccharide is bound at subsites +1 to +4 of the reducing end (aglycon portion).
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